Key
to subfamilies, tribes and genera in Costa Rica and Panama |
| 1 |
Antennal funicle of four articles;
tarsus of five distinct articles; size small, total body length less
than 4 mm |
Dryophthorinae |
Dryophthorus Germar, 1824
and Stenommatus Wollaston, 1873 |
| |
- Antennal funicle of six articles;
tarsus of five articles, but article 4 small and difficult to see
at base of article 3; size small to large, total body length greater
than 3 mm and generally greater than 10 mm |
2 |
 |
| 2 |
Front coxae contiguous; anterior margin of pronotum
with postocular lobe; pygidium largely covered by elytra; tarsus with
article 3 subequal in width to article 2; mandibles triangular, directed
anteriorly and with inner edge lacking teeth |
Orthognathinae, Orthognathini |
3 |
| |
- Front coxae distinctly (but in some
taxa, narrowly) separated; anterior margin of pronotum straight behind
eye; pygidium broadly exposed beyond elytra (most) or largely covered
by elytra (few); tarsus with article 3 noticeably wider than article
2 and generally bilobed or occasionally subequal in width to article
4; mandibles variously shaped |
4 |
 |
| 3 |
Front tibia with outer apical angle
produced anterolaterally to a sharp or narrowly rounded angle; middle
and hind legs robust, tibiae much wider at apex than at base, fossorial
in form |
Orthognathus Schoenherr, 1838 |
| |
- Front tibia with outer apical angle
not produced anterolaterally, broadly rounded; middle and hind legs
long and narrow, tibiae with apex and base subequal in width or with
apex slightly wider than base, not fossorial in form |
Mesocordylus Lacordaire, 1866 |
 |
| 4 |
Size small, total body length less
than 5 mm; tibia with distinct subapical tooth at inner angle in addition
to larger uncus |
5 |
| |
- Size small to large, total body length
greater than 5 mm and generally greater than 10 mm; tibia with either
a small rounded subapical swelling (most) or sharp subapical tooth
(few) at inner angle in addition to larger uncus |
6 |
 |
| 5 |
Rostrum straight, in lateral view with base continuous
with head; eyes clearly visible in dorsal view |
Rhynchophorinae, Litosomini (part) |
Sitophilus Schoenherr, 1838 |
| |
- Rostrum curved ventrally, in lateral view separated
from head by basal constriction; eyes not or barely visible in dorsal
view |
Rhynchophorinae, Polytini |
Polytus Faust, 1894 |
 |
| 6 |
Mandibles large, divaricate, divergent, with distinct
teeth on outer edge; outer face concave medially; pygidium largely
covered by elytra |
Orthognathinae, Rhinostomini |
Rhinostomus Rafinesque, 1815 |
| |
- Mandibles small, convergent and occluding
medially, with inner margins distinctly toothed (most); or, elongate
subtriangular, with inner margins parallel-sided and lacking teeth,
not touching medially (few); pygidium broadly exposed beyond elytra |
7 |
 |
| 7 |
Size very large, total body length greater than
25 mm; metepisternum very broad, length more or less 2 times width |
Rhynchophorinae, Rhynchophorini |
8 |
| |
- Size small to large, total body length
greater than 5 mm but usually less than 25 mm; metepisternum narrow,
length 3 or more times width |
9 |
 |
| 8 |
Scutellum large, with elongate apical
extension; rostrum glabrous in female, with elongate, dorsal patch
of long, dark brown pilosity in male |
Rhynchophorus Herbst, 1795 |
| |
- Scutellum small, apically acuminate
but not extended; rostrum glabrous in both sexes |
Dynamis Chevrolat, 1883 |
 |
| 9 |
Rostrum extremely long and slender, as long as
pronotum and elytra combined; mandibles elongate subtriangular, with
inner margins parallel-sided and lacking teeth, not touching medially;
antennal club glabrous basally, with slender, stalk-like base, width
at apex 3-4 times width at base; legs long and slender |
Rhynchophorinae, Litosomini (part) |
Toxorhinus Lacordaire, 1866 |
| |
- Rostrum short and robust to long
and slender, but shorter than pronotum and elytra combined; mandibles
small, convergent and occluding medially, with inner margins distinctly
toothed; antennal club with glabrous broad base, width at apex less
than 3 times width at base; legs robust |
10 |
 |
| 10 |
Body cylindrical; pronotum very slightly shorter
than elytra; middle and hind tibiae short, straight, slightly shorter
than combined length of tarsal articles 1-4; tibial apices with three
distinct apical projections, two (one long, one short) at inner angle,
one (short) at outer angle; elytra with distinct long erect setae;
interval 2 densely pilose throughout most of length |
Rhynchophorinae, Sphenophorini (part) |
Alloscolytoproctus Hustache, 1929 |
| |
- Body somewhat to markedly dorsoventrally
compressed; pronotum distinctly shorter than elytra; middle and hind
tibiae long, straight or curved, longer than combined length of tarsal
articles 1-4; tibial apices with single uncus at inner angle (in some
also with small rounded subapical swelling or tooth); elytra without
long erect setae; no intervals densely pilose |
11 |
 |
| 11 |
Scutellum (exposed portion) widest at or near
middle, shape rhomboidal or subcircular, more or less as long as wide |
Rhynchophorinae, Litosomini (part) |
12 |
| |
- Scutellum (exposed portion) widest at or near
base, shape triangular or subtriangular, generally longer than wide |
Rhynchophorinae, Sphenophorini (part) |
16 |
 |
| 12 |
Rostrum markedly laterally compressed
in at least apical one-half; body flat, maximum width greater than
1.35 times depth |
Melchus Lacordaire, 1866 |
| |
- Rostrum more or less cylindrical,
not laterally compressed; body thicker, maximum width less than 1.35
times depth |
13 |
 |
| 13 |
Elytra with scattered concentrations
(nodules) of short erect setae, in some cases surrounded by dense
micropilosity |
14 |
| |
- Elytra glabrous or with at most scattered,
sparse setae or scales |
15 |
 |
| 14 |
Rostrum curved ventrally; antennal
club with apical pilose part one-third to one-half length of basal
glabrous part; pronotum and/or elytra with scattered patches (nodules)
of micropilosity; front femur of male with distinct subapical swelling
or tooth on inner margin |
Daisya Anderson, 2003 |
| |
- Rostrum straight; antennal club with
apical pilose part obliquely oriented and appearing only as a narrow
line; entire body uniformly covered with brownish matte coating; front
femur of male without subapical swelling or tooth on inner margin |
Neophrynoides O’Brien and Wibmer, 1982 |
 |
| 15 |
Rostrum curved ventrally; elytral
striae impressed, intervals with single row of fine punctures; pronotum
uniformly punctate |
Cosmopolites Chevrolat, 1885 |
| |
- Rostrum more or less straight; elytral
striae very shallowly, indistinctly impressed, intervals not punctate;
pronotum virtually impunctate |
Eucalandra Faust, 1899 |
 |
| 16 |
Tarsal article 3 with long ventral
pilosity, confined to apical margin as a continuous fringe, ventral
surface otherwise glabrous; antennal club obliquely truncate at apex
with apical pilose part very short, appearing recessed within glabrous
part, visible only as a narrow line in lateral view |
Scyphophorus Schoenherr, 1838 |
| |
- Tarsal article 3 with various ventral
pilosity; long or short, uniformly covering 1/3 or more of ventral
surface, or with pilosity sparse and confined to anterolateral angle
or lateral margins, ventral surface otherwise glabrous; antennal club
various, with apex evenly (most) or obliquely (few) truncate; apical
pilose part of club short or long, generally visible as more than
a narrow line in lateral view |
17 |
 |
| 17 |
Tarsal article 3 with ventral pilosity
confined to anterolateral angle or lateral margins |
Sphenophorus Schoenherr, 1838 |
| |
- Tarsal article 3 with ventral pilosity
more extensive, covering 1/3 or more of ventral surface |
18 |
 |
| 18 |
Tarsal article 5 (claw-bearing article)
with venter at apex excavate or markedly to slightly bilamellate (difficult
to see in some specimens); antennal scape enlarged and laterally compressed
(few) or more or less cylindrical (most); rostrum humped with abrupt
change of orientation from anteriorly directed near base to ventrally
directed near apex (many) or anteroventrally oriented, without any
dorsal hump or abrupt change in orientation (many); legs of males
and females generally similar in shape and vestiture; pygidium narrow
especially towards apex which generally is narrowly rounded, tumescent
subapically and extended beyond apex of ventrite 5 in ventral view;
male aedeagus without lateral line |
Rhodobaenus LeConte, 1876 (in part) |
| |
- Tarsal article 5 (claw-bearing article)
with apical venter smooth; antennal scape more or less cylindrical
(most) or laterally compressed (few); rostrum generally uniformly
anteriorly or anteroventrally oriented, without any dorsal hump or
abrupt change in orientation (most) or rostrum humped with abrupt
change in orientation from anteriorly directed basally to ventrally
directed apically (few); legs of males and females either generally
similar in structure (many) or with front or hind tibiae of males
of a different shape, or bearing different vestiture in the form of
rows or tufts of longer hairs (many); pygidium broad especially towards
apex which generally is broadly truncate, flat or variously carinate
subapically and at most slightly extended beyond apex of ventrite
5 in ventral view; male aedeagus with or without lateral line |
19 |
 |
| 19 |
Male aedeagus with lateral line (complete
or incomplete); legs with front or hind tibiae of males bearing vestiture
in the form of rows or tufts of longer hairs (which are not present
in females) or of a different shape (most), or legs of both sexes
generally similar in shape and vestiture (few); ventrally, lacking
tubercles between front coxae or with metasternum more or less flat
between mesocoxae |
Metamasius Horn, 1876 |
| |
- Male aedeagus without lateral line;
legs of males and females generally similar in shape and vestiture;
ventrally, some species with variously developed tubercles between
front coxae or with metasternum markedly produced and subconical between
mesocoxae |
20 |
 |
| 20 |
Small (8.0-13.0 mm; few) to large
(13.0-22.0 mm, most) entirely grey or black species, some with dark
velvety black markings or a pale colored "V" or "C" shaped marking
on the elytra; ventrally, lacking tubercles between front coxae or
with metasternum more or less flat between mesocoxae |
Rhodobaenus LeConte, 1876 (in part) |
| |
- Small (7.0-8.0 mm; few) to large
(12.0-25.0 mm, most) black and red, orange or yellow species (most),
or entirely black but then metasternum markedly produced and subconical
between mesocoxae (few) or pronotum and elytra distinctly, irregularly
sculptured (few); ventrally, with variously developed tubercles between
front coxae (many) or with metasternum markedly produced and subconical
between mesocoxae (many) |
Cactophagus LeConte, 1876 |