CARABIDAE
Author: John F. Lawrence.
Classification
Suborder Adephaga
Description
Antenae with 11 segments, filiform or rarely perfoliate. Antennal insertions exposed or concealed. Visible portion of procoxa globular with the trochantin concealed. Procoxal cavity externally open to closed and internally open to closed. Mesocoxae separated by less than 0.4 X coxal width to 1 coxal width, with mesocoxal cavity laterally open to closed. Tarsal formula 5-5-5. Numberof ventrites 6 with 3 connate. Body length 1-40 mm. Variable form and vestiture. Antennae perfoliate only in Homopterus. Metacoxal plates developed mesally only. Usually with distinct antenna cleaner on protibia. Prominent tactile setae on upper surface.
Quick identification
Carabidae contains the majority of terrestrial Adephaga and most are easily identified using characters such as the filiform antennae with antennomeres beyond 3 or 4 densely pubescent, a grooming organ or antenna cleaner on the protibia, presence of erect, fixed tactile setae on the body. There are a number of unusual carabids which might be misplaced. Some of the Paussinae lack an antenna cleaner and tactile setae, and may have highly modified antennae (first figure). The genus Omophron (2nd figure) has an atypical shape for the group and looks like some of the aquatic Adephaga. Another group of unusual carabids is the subfamily Pseudomorphinae (fourth figure) which are broad and flattened.
Genera occurring in Costa Rica
Paussinae: Goniotropis, Homopterus, Ozaena, Pachyteles, Physea, Platycerozaena...
Omophroninae: Omophron.
Carabinae: Calosoma, Notiophilus.
Cicindelinae: Cicindela, Ctenostoma, Iresia, Megacephala, Odontocheila, Oxycheila, Pseudoxycheila...
Scaritinae: Ardistomis, Aspidoglossa, Clivina, Dyschirius, Oxydrepanus, Pasimachus, Scarites, Schizogenius...
Trechinae: Bembidion, Cnides, Elaphropus, Micratopus, Mioptachys, Paratachys, Paratrechus, Pericompsus, Tachyta, Tachys...
Harpalinae: Abaris, Agra, Alkestis, Amara, Anatrichis, Anchonoderus, Ancystroglossus, Anisocnemus, Apenes, Apristus, Badister, Barysomus, Bradycellus, Calleida, Calophaena, Calybe, Catapiesis, Catascopus, Colliuris, Coptia, Coptodera, Dercylus, Dromius, Ega, Epikastea, Eucheila, Eurycoleus, Galerita, Galerucidia, Helluobrochus, Helluomorphoides, Homalomorpha, Inna, Lachnophorus, Lebia, Leptotrachelon, Leptotrachelus, Loxandrus, Morion, Moriosomus, Nemotarsus, Notiobia, Onypterygia, Oxycrepis, Panagaeus, Pelecium, Pelmatellus, Pentagonica, Perigona, Pinacodera, Platynus, Pseudaptinus, Selenophorus, Stenocrepis, Stenognathus, Stenolophus, Stenomorphus, Tetragonoderus, Zuphium...
Pseudomorphinae: Pseudomorpha.
Brachininae: Brachinus, Pheropsophus.
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