Janzen, D.H. 1980. Specificity of seed-attacking beetles in a Costa Rican deciduous forest. Journal of Ecology, 68:929-952.

Kingsolver, J.M. 1980. Eighteen new species of Bruchidae principally from Costa Rica with host records and distributional notes (Insecta: Coleoptera). Proc. Biol. Soc. Washington, 93:229-283.